Tout_velu -> dally_like
Tout_velu -> division_abnormally_delayed
dally_like -> Hh_exterior_input
division_abnormally_delayed -> Hh_exterior_input
Rasp -> Hh_exterior_input
Shifted -> Hh_exterior_input
interference_hedgehog -> Hh_exterior_input
Brother_of_iHog -> Hh_exterior_input
Dispatched -> Hh_exterior_input
Lipophorin -> Hh_exterior_input
Hh_exterior_input -| Pka
Pka -| Smoothened
G_protein_receptor_coupled_kinase_2 -> Smoothened
Target_of_Wingless -| Smoothened
Smoothened -> Fused
Smoothened -| Costa
Fused -| Costa
Fused -| Suppressor_of_Fused
Costa -| Ci_activator
Suppressor_of_Fused -| Ci_activator
Shaggy -| Ci_activator
Protein_kinase_A -| Ci_activator
Supernumary_limbs -| Ci_activator
Casein_kinase_1_alpha -| Ci_activator
Costa -> Ci_repressor
Suppressor_of_Fused -> Ci_repressor
Shaggy -> Ci_repressor
Protein_kinase_A -> Ci_repressor
Supernumary_limbs -> Ci_repressor
Casein_kinase_1_alpha -> Ci_repressor
Ci_activator -> Targets
Ci_repressor -| Targets
$Ci_activator: (((((!Costa | (Costa & !Suppressor_of_Fused)) | ((((Costa & Suppressor_of_Fused) & !Shaggy) & !Protein_kinase_A) & !Supernumary_limbs)) | (((((Costa & Suppressor_of_Fused) & !Shaggy) & !Protein_kinase_A) & Supernumary_limbs) & !Casein_kinase_1_alpha)) | ((((Costa & Suppressor_of_Fused) & !Shaggy) & Protein_kinase_A) & !Supernumary_limbs)) | (((Costa & Suppressor_of_Fused) & Shaggy) & !Supernumary_limbs))
$Ci_repressor: (((((Costa & Suppressor_of_Fused) & Shaggy) & Protein_kinase_A) & Supernumary_limbs) & Casein_kinase_1_alpha)
$Costa: (!Smoothened & !Fused)
$Fused: Smoothened
$Hh_exterior_input: (((((((dally_like & division_abnormally_delayed) & Rasp) & Shifted) & interference_hedgehog) & Brother_of_iHog) & Dispatched) & Lipophorin)
$Pka: !Hh_exterior_input
$Smoothened: ((!Pka & G_protein_receptor_coupled_kinase_2) & !Target_of_Wingless)
$Suppressor_of_Fused: !Fused
$Targets: (Ci_activator & !Ci_repressor)
$dally_like: Tout_velu
$division_abnormally_delayed: Tout_velu